Distribution
Africa: Ruanda-Urundi rivers, rivers of eastern highlands of Zimbabwe and the Transvaal escarpment (Ref. 2801), Cunene, Cuanza, Okavango, Zambezi, east coast systems south to the Incomati (Komati) system and Lake Tanganyika (Ref. 52193). In the Congo system in the the upper Lualaba, Luapula (Ref. 41590), Lake Mweru (Ref. 11970) (upper Congo River basin) and upper Kasai (Ref. 41590).