Distribution
Africa: Angola, Botswana, Namibia, Zambia, Zimbabwe and Democratic Republic of the Congo in the Cunene River system (Angola and Namibia); Okovango River and swamps; upper Zambezi system including Lake Cameia, Kafue River and Luangwa system; Luapula-Mweru, Lulua River and Angolan Kasai (upper and middle Congo River basin).