Distribution
North America: Barranca de Venados, Puente Venado, R?o de Metztitl?n, Hidalgo, Mexico (type locality) (Ref. 33021). South of the R?o P?nuco in five coastal rivers Tuxpan, Cazones, Tecolutla, Nautla and Misantla rivers) of the eastern stretch of the Trans-Mexican Volcanic Belt (TMB) at 20?N (Ref. 80389).