Distribution
Africa: Senegal, Gambia, Casamance, Corubal, Geba, Little Scarcies, Sassandra, Boubo, Bandama, Agn?bi, M?, Como?, Tano, Pra, Volta basin, Mono, Sio, Ou?m?, Ogun, entire Niger and Chad basins, Cross, Wouri and Sanaga; absent between Little Scarcies and Sassandra; further in the entire Congo River basin, Malagarasi, Luiche, Cunene, Okavango, entire Zambezi including the Kafue system, Lake Kariba, Webi Shebeli, Wando, Tana, Galana, Athi, Wami, Rufiji-Ruaha, Lake Rukwa, lower Sabi-Lundi, Shire below Murchison Falls, Lake Chilwa, Ruvu, Pungwe, Limpopo, Incomati and Pongola (Ref. 43912).