Distribution
Europe: Great Britain north to Scottish highlands, rivers draining to North Sea north to Scotland and about Stavanger , Baltic Sea basin, Atlantic as far south as Adour drainage and isolated populations in Sado, Tagus and Douro drainages , Mediterranean basin in France and western Italy. Locally in Ireland, upper Volga, upper Danube and some of their tributaries and Pescara drainage on Adriatic coast of Italy. Appendix III of the Bern Convention.