Distribution
Europe: Baltic: Bothnian Gulf, Gotland (missing along Swedish coast southward of Stockholm); southern Baltic: from Germany to Gulf of Finland; several landlocked populations in Sweden (Lakes V?nern, V?ttern, Locknesj?n, Storvindeln, Storlaisan, Skalka-Parkijaur, Arjeplog) and Russia (Neva drainage, Lakes Ladoga and Onega).