Distribution
Africa: Zambezi, Kafue and Shire Rivers; Upper Congo system; Lakes Tanganyika, Bangweulu, Kobo, Niumbe, Mweru and Malawi; Chobe, Okavango and Cunene Rivers; Pungwe, Sabi, Lundi Rivers and Zimbabwean tributaries of the Limpopo River; Incomati, Pongolo and Umgeni rivers and Lake Sibaya in Natal.