Distribution
Africa: Lake Victoria, Edward, George, small lakes of Uganda, Lake Kivu, smaller lakes of Rwanda, Lake Tanganyika, Bangweulu-Moero system, in Malawi, Kagera, Malagarazi, Ruzizi and Tana Rivers. Cunene, Okavango, upper Zambezi, and Kafue systems, Zambian Congo, Central Africa Great Lakes (Ref. 7248).